"sample" "PC1: 48% variance" "PC2: 24% variance" "gonad" -162.927921718451 -102.309686813727 "heart" -157.585981677597 -48.3813646744006 "ctenidia" -82.9843909215941 113.88534782287 "juvenile" 129.816510723806 173.187548293266 "larvae" 273.681783593837 -136.381844628009